This new TV special tells the two years of Luffy's training with Silvers Rayleigh in Rusukaina Island after the Battle of Marineford, which has never been told in the manga or TV anime. It also features a new villain named Burndy World who is known as "The Destroyer of the World" and has the power of the MoaMoa Fruit (its ability is still unknown). Luffy has to protect Boa Hancock from the new enemy who escaped from the level 6 of Impel Down. He is newly designed by the original creator of the manga, Eiichiro Oda.

The special takes place during the two year before the Straw Hats reunite on Sabaody. Luffy is currently in Rusukaina training to get stronger to take on the New World. However the training is interrupted when Hancock's sisters, Marigold and Sandersonia, are kidnapped by the Byrnndi World, a pirate who was locked away on Level 6 of Impel Down but escaped during Luffy's invasion to save Ace, in order to lure Hancock to him and use her as a hostage against the World Government due to her Shichibukai status. Thus Luffy and Hancock head off to confront him and save Hancock's sisters.

Zephyr, now known as Z, rides the seas with only one goal: Destroy all pirates and their dreams at becoming King of Pirates. When Luffy and his crew encounter him at sea, not only are they utterly defeated by the man with an arm made of Seastone, Nami, Robin, and Chopper are turned 10 years younger due to Z's minion Ain. Luffy is so determined to win against him that he does not even notice Z's master plan that could sacrifice thousands of lives.

The Straw Hat Pirates are taking on Gild Tesoro, one of the richest men in the world.

Monkey D. Luffy and his Straw Hat Crew have finally arrived on Gran Tesoro, a ship carrying the largest entertainment city in the world. Drawn in by the chances of hitting the jackpot, the crew immediately head to the casino. There, they quickly find themselves on a winning streak, playing with what seems to be endless luck. When offered a special gamble by Gild Tesoro—the master of the city himself—the crew agrees, choosing to believe in their captain's luck. However, when they find themselves victims of a despicable scam, the crew quickly realize that there is something darker happening beneath the city's surface. Left penniless and beaten down, the Straw Hat Crew are forced to rely on another gamble of a plan. With the help of a new friend or two, the group must work to reclaim what they've lost before time, and what remains of their luck, runs out.

A short prequel to One Piece Film: Gold featuring the nine Straw hat crew members on the ship, planning what to do when they arrive the Casino.

Upon hearing news that islands in East Blue are being destroyed, Monkey D. Luffy and his crew go to investigate. On their way, however, an outlandish pirate ship appears out of the sky, helmed by the infamous pirate Shiki "the Golden Lion"—a man who ate the Float-Float Fruit and the first ever prisoner to escape from Impel Down. In his quest to defeat the World Government, Shiki kidnaps Nami to be his own navigator and sends the rest of the Straw Hat Pirates to his floating islands as hostages, leaving her in a dilemma. Separated in a land under Shiki's absolute control, Luffy and his crew must survive the mystifying terrain in order to bring back their navigator and friend.

The Straw Hat Pirates enter the rough seas of the New World in search of the hidden treasures of the Pirate King, Gol D. Roger－One Piece. On their voyage, the pirates come across a terrifying, powerful man, former Marine Admiral Z. Z is accused of having stolen the "Dyna Stones", weapons believed to have the power to shake up the New World. The Marine Headquarters believes Z is about to use it to end the pirate era, and with it, the lives of many innocent people. In fear of such a phenomenal event, marines start to take action against the former admiral. Even if it means stumbling upon marines and the navy, the Straw Hat Pirates decided to chase after Z and stop him from causing havoc. As they continue to embark on their ventures, the pirates bump into new and familiar acquaintances.

The story of the Hand Island no Bouken special is set a little before the events of One Piece Film Z and depicts a major "Hand Island" incident during the New World storyline. The story is set at "Cannon Town" where a parent and a child work as craftspeople. The incident revolves around them and a commodore of a marine base. The special will also have a flashback scene to the story in the first chapter of the manga, but with new animation.

There once was a pirate known as the Great Gold Pirate Woonan, who obtained almost 1/3 of the world's gold. Over the course of a few years, the pirate's existence faded, and a legend grew that he disappeared with his gold to a remote island, an island pirates continue to search for. Aboard the Going Merry, Luffy and his crew, starved and reckless, are robbed of their treasure. In an attempt to get it back, they wreck the getaway ship, guided by a young boy named Tabio, who's a captured part of El Drago's pirate crew. El Drago's love for gold has driven him to look for Woonan's island, and thanks to Woonan's treasure map, he finds it. During this time, Luffy's crew have been split up, and despite their own circumstances, they must find a way to stop El Drago from obtaining Woonan's gold.

20 years after his escape from Impel Down, the legendary pirate Shiki, the Golden Lion, reappears causing massive upheaval to the Marines. During his long seclusion, he was able to come up with a scheme to bring the World Government to his knees. On his way to execute the plan, Shiki crosses paths with the Straw Hat Pirates and becomes so impressed with Nami's knowledge of meteorology that he abducts her to forcedly enlist her into his crew. Luffy and the gang end up on a strange land populated with monstrous beasts as they desperately search for Shiki and Nami.

The world's greatest exposition of the pirates, by the pirates, for the pirates—the Pirates Festival. Luffy and the rest of the Straw Hat Crew receive an invitation from its host Buena Festa who is known as the Master of Festivities. They arrive to find a venue packed with glamorous pavilions and many pirates including the ones from the Worst Generation. The place is electric.